The freshwater and terrestrial turtles from Monte Pila and Fuenmayor (La Rioja, northern Spain): new data on the lower Miocene turtle diversity of the Ebro Basin

Abundant turtle fossils from two lower Miocene localities in which this lineage had not previously been studied are described herein. The Monte Pila and Fuenmayor, 10 km apart, both located the Autonomous Community of La Rioja (northern Spain), Ebro Basin. Information about turtles levels basin was until now restricted to finds made MN3 biozone Bardenas Reales Navarre. Several taxa recognized there. However, material attributed each them scarce. Fuenmayor MN2. remains abundant sites. A terrestrial lineage, at Pila, is represented fauna It represents first record TitanochelonPérez-García & Vlachos, 2014 one oldest member genus, being assigned Titanochelon cf. bolivari (Hernández-Pacheco, 1917). only identified most Ptychogasterinae. All clade freshwater there compatible with a single form, for high range intraspecific variability recognized. Ptychogaster (Temnoclemmys) bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola Astibia, 1999, species having recognized, now, as exclusive